When a murder of a physicist, Leonardo Vetra, finds a symbolist, Robert Langdon, and Mr. Vetra's daughter, Vittoria, on an adventure for a secret brotherhood, The Illuminati. Clues lead them all around the Vatican, including the four alters of science, Earth, Air, Fire and Water. An Assassin, working for the Illuminati, has captured four cardinals, and murders each, painfully. Robert and Vittoria also are searching for a new very destructive weapon that could kill millions.

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When the Museum of Natural History is closed for upgrades and renovations, the museum pieces are moved into federal storage at the famous Washington Museums. The centerpiece of the film will be bringing to life the Smithsonian Institution, which houses the world's largest museum complex with more than 136 million items in its collections, ranging from the plane Amelia Earhart flew on her non-stop solo flight across the Atlantic and Al Capone's rap sheet and mug shot to Dorothy's ruby slippers, Fonzie's jacket from Happy Days, the still from M*A*S*H and Archie Bunker's lounge chair from All in the Family.